Ciencia · 2012
The genus Eragrostis (Poaceae: Chloridoideae) in northwestern South America (Colombia, Ecuador, and Peru): morphological and taxonomic studies
A morphological and taxonomic treatment of Eragrostis for northwest South America is given. The present revision intends to clarify the taxonomic and nomenclatural problems of the species of Eragrostis present in northwest South America. The classification, nomenclature, affinities, and macro and micromorphological variation of the genus are discussed. Forty-five morphological and micromorphological characters were selected for use in the descriptions and differentiation among the species. This study is based on the examination of herbarium specimens from AAU, AMAZ, ANSM, BA, BAA, BAB, BM, CA, CAUP, COAH, COL, CORD, CPUN, CTES, CUZ, ENCB, F, GB, GH, HAO, HUA, HUT, K, LE, LIL, LINN, LP, MA, MEXU, MICH, MO, NY, P, PSO, QCA, RSA, S, SGO, SI, TAES, UC, UPS, US, USM, UTC, W, and WIS, including the type specimens of most species studied. In addition, extensive field studies were performed to assess the diversity of the genus in this region of South America. Thirty-seven species are recognized in the study area (24 native and 13 introduced-naturalized); four of the species are cited for the first time for Colombia [Eragrostis atrovirens (Desf.) Trin. ex Steud., Eragrostis barrelieri Daveau, Eragrostis gangetica (Roxb.) Steud., Eragrostis soratensis Jedwabn.] and three for Peru [Eragrostis acutiflora (Kunth) Nees, Eragrostis bahiensis Schrad. ex Schult., and Eragrostis secundiflora J. Presl]. Eragrostis condensata (J. Presl) Steud. is endemic to Ecuador, while Eragrostis magna Hitchc. and Eragrostis pilgeri Fedde are endemic to Peru. Twenty-nine species are recorded in Colombia, 19 in Ecuador, and 26 in Peru. Keys for determining the species, descriptions, illustrations, geographical distributions, morphological and ecological observations, chromosome numbers, specimens studied, and synonymies are provided for all native and adventive species of Eragrostis in northwestern South America. Eragrostis lasseri Luces is placed as a synonym of Eragrostis ciliaris (L.) R. Br., and Eragrostis nigricans (Kunth) Steud. var. punensis Nicora is placed as a synonym of Eragrostis mexicana (Hornem.) Link subsp. mexicana. The lectotype for Eragrostis barrelieri Daveau is designated here. Eragrostis patula (Kunth) Steud. and Eragrostis unioloides (Retz.) Nees ex Steud. are excluded from the Colombian Flora since all specimens identified as Eragrostis patula are apparently misidentified and are usually referable to Eragrostis tenuifolia (A. Rich.) Hochst. ex Steud. A single specimen identified as Eragrostis unioloides as cited in the Flora of Antioquia (Colombia) was misidentified and it belongs to Chascolytrum juergensii (Hack.) Essi, Souza-Chies & Longhi-Wagner. (texto tomado de la fuente)
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Contenido
- Macromorphological analysesp. 23
- Micromorphological analysesp. 23
- Phytolithsp. 27
- Reproductive structures23
- Microscopy23
- Micromorphological and anatomical terminology24
- Species concept24
- Ant samples24
- Phytolithsp. 27
- Silica bodies27
- Long cells27
- Short cells27
- Stomatal complexes27
- Macrohairs27
- Microhairs27
- Prickle hairs27
- Papillae27
- Glandsp. 27
- An overview of glands in grasses27
- Microhairs (bicellular trichomes) in Eragrostis31
- Multicellular glands (extrafloral nectaries) in Eragrostis33
- Spikelet morphologyp. 39
- Caryopsis morphologyp. 45
- Diversity of species of Eragrostis in northwestern South Americap. 56
- Ecology and geographic distributionp. 57
- Economic importancep. 58
- Taxonomic treatmentp. 59
- Eragrostis Wolf61
- Etymology61
- Base chromosome number and levels of ploidy61
- Monophyly of Eragrostis62
- Biochemical features62
- The subtribe Eragrostidinae62
- Key to separate Eragrostis and Steirachnep. 63
- Infrageneric classification63
- Commentsp. 1
- Key to the species of Eragrostis in Colombiap. 67
- Key to the species of Eragrostis in Ecuadorp. 73
- Key to the species of Eragrostis in Perup. 79
- Descriptions of the speciesp. 83
- Eragrostis acutiflora (Kunth) Neesp. 33
- Eragrostis airoides Neesp. 33
- Eragrostis andicola R. E. Frp. 56
- Eragrostis atrovirens (Desf.) Trin. ex Steudp. 33
- Eragrostis attenuata Hitchcp. 56
- Eragrostis bahiensis Schrad. Ex Schultp. 33
- Eragrostis barrelieri Daveaup. 33
- Eragrostis cilianensis (All.) Vignolo ex Janchp. 33
- Eragrostis ciliaris (L.) R. Brp. 33
- Eragrostis condensata (J. Presl) Steudp. 56
- Eragrostis curvula (Schrad.) Neesp. 33
- Eragrostis gangetica (Roxb.) Steudp. 33
- Eragrostis hypnoides (Lam.) Britton, Sterns & Poggenbp. 188
- Eragrostis intermedia Hitchcp. 56
- Eragrostis japonica (Thunb.) Trinp. 56
- Eragrostis lugens Neesp. 47
- Eragrostis lurida J. Preslp. 47
- Eragrostis magna Hitchcp. 47
- Eragrostis maypurensis (Kunth) Steudp. 56
- Eragrostis mexicana (Hornem.) Linkp. 33
- Eragrostis mokensis Pilgp. 48
- Eragrostis nigricans (Kunth) Steudp. 56
- Eragrostis pastoensis (Kunth) Trinp. 56
- Eragrostis pectinacea (Michx.) Neesp. 33
- Eragrostis peruviana (Jacq.) Trinp. 57
- Eragrostis pilgeri Feddep. 48
- Eragrostis pilosa (L.) P. Beauvp. 33
- Eragrostis polytricha Neesp. 48
- Eragrostis prolifera (Sw.) Steudp. 33
- Eragrostis rufescens Schrad. ex Schultp. 57
- Eragrostis secundiflora J. Preslp. 57
- Eragrostis soratensis Jedwabnp. 57
- Eragrostis tenella (L.) P. Beauv. ex Roem. & Schultp. 33
- Eragrostis tenuifolia (A. Rich.) Hochst. ex Steudp. 33
- Eragrostis unioloides (Retz.) Nees ex Steudp. 33
- Eragrostis viscosa (Retz.) Trinp. 57
- Eragrostis weberbaueri Pilgp. 57
- Excluded namesp. 165